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From this book: If generations overlap, then the effective population size $N_e$ does not equal the population size $N$.

I know mathematical formulations in order to find the effective population size $N_e$ when the sex-ratio is biased $\left(N_e = \frac{4N_mN_f}{N_m+N_f}\right)$ or when the population size varies cyclically through time $\left(N_e = \frac{n}{\sum_{i=1}^n\frac{1}{N_i}}\right)$. Similarly, how can one calculate the effective population size in a population with overlapping generations?

To answer this question one will typically need to use variables such as $T_G$ the generation time and $T_R$ the time between two reproductive event (or only their ratio).

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Some info on this can be found in Hendrick (2010, p 234)‌​, and if you follow the reference there (Laikre et al, 1998) you might find a more complete answer. –  fileunderwater Mar 18 at 11:26

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