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How do you define a male-biased gene and a female-biased gene as they are found in the abstract of this article.

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perhaps by strong difference in expression between two sexes. –  WYSIWYG May 23 at 10:12
    
WYSIWYG yes, that's what I was thinking too. Let's wait and see if someone can make sure of this. –  Remi.b May 24 at 8:09

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up vote 1 down vote accepted

In the linked paper, the authors discuss this as sex-based gene expression that evolved by sex-specific selection. The expression is not limited to one sex (which are sex-limited genes). Sex-biased genes are expressed by both sexes, but differently between sexes.

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