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A girl pretended having a piece of food in her hand and pointed it to a Barbary Macaque in Gibraltar. As a response he made this facial expression to her. What does this mean?

GibraltarBarbaryMacaqueExpressions

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What was the sound that it made? Macaque vocalizations are also well studied, and it looks like it was vocalizing. –  Atl LED Aug 13 at 13:20
    
@Atl LED The macaque didn't make any sound, its lips were maybe more like this: upload.wikimedia.org/wikipedia/commons/e/e1/… and its eyes wide open –  Mehdi Nellen Aug 14 at 11:39

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That is a threat face. Barbary macaque threat faces often appear with a brow raise, lowered head, and an o-shaped mouth, sometimes with and sometimes without a vocalization. Given the context you described it is not surprising the girl received a threat. *Based on personal research experience

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It would be fantastic if you provided some references. –  Bez Aug 14 at 18:26

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