The application of dynamic systems and game theory to evolving populations in biology, economics, and social sciences. It is particularly well suited for studying social dilemmas like the evolution of cooperation and altruism.

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Queller's 1985 version of hamilton's rule

Queller 1985 ("Kinship, reciprocity and synergism in the evolution of social behavior") provides a generalization of Hamilton's rule that allows for non-additivity. To accomplish that, Queller writes ...
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Why does a pecking order almost always exist in groups of social mammals?

I think the best explanation might come from evolutionary game theory. But, an another viewpoint may be equally important.