Two cytoplasmic genomes exist in plant in addition to the nuclear one. As far as I understand, they divide more or less during the entire life of the plant along with cell proliferation/growth in meristem. That means that somatic mutations arise. Can this variation accumulate and be passed on to offsprings? Let's say there is a mutation in plastid DNA that spread in a meristem before flower formation, will you find these mutation in the pollen/ovule plastid ? Or is there a way to keep the original genome intact for germinal cell? e.g: stock of proplastid/mitochondria in meristems that doesn't divide before reproductive tissue formation?