From a 2017 paper that suggests the more distant (and common) origin of both HCoV-229E and HCoV-NL63 is in African bat coronaviruses, I see that the closest virus most similar to HCoV-229E is Aplaca-CoV (Alpaca is a camelid native to South America.)
Phylogenetic relationships of NL63-like and 229E-like viruses at the receptor binding domain of HCoV-NL63. The tree is based on an amino acid alignment and midpoint rooted.
Am I reading this correctly? Is Alpaca-CoV still the closest known (animal) relative of HCoV-229E? (Or is that just at the RBD level?)