Hopefully this syllogism will answer your question.

Given the following premises:

1. In the absence of selection, fitness of individuals are at a theoretical maximum.
2. If a theoretical maximum fitness is achieved the effective population size is also at a maximum.
3. If there is an allele that confers both increased fitness and decreased fitness (simultaneously) you have a genetic conflict (e.g. an allele that turns you green gives you good woodland camouflage but is un-attractive).
4. If a deleterious allele is linked to a beneficial allele this is similar to having a genetic conflict (mentioned in 3.).

The conclusion follows:

Strong linkage between beneficial alleles and deleterious alleles (Hill Robertson effects, i.e. 3. and 4.) prevents a theoretically maximum fitness to be attained (1.) and thus a reduced effective population size (2.). Which can only be overcome by high levels of recombination and mating (i.e. by separating deleterious and beneficial alleles).

Hopefully this syllogism will answer your question.

Given the following premises:

1. In the absence of selection, fitness of individuals are at a theoretical maximum.
2. If a theoretical maximum fitness is achieved then effective population size is maximum.
3. If there is an allele that confers both increased and decreased fitness you have a genetic conflict (e.g. an allele that turns you green gives you good woodland camouflage but is un-attractive).
4. If a deleterious allele is linked to a beneficial allele this is similar to having a genetic conflict.

The conclusion follows:

Strong linkage between beneficial alleles and deleterious alleles (Hill Robertson effects, i.e. 3. and 4.) prevents a theoretically maximum fitness to be attained (1.) and thus a reduced effective population size (2.). Which can only be overcome by high levels of recombination and mating (i.e. by separating deleterious and beneficial alleles).

Hopefully this syllogism will answer your question.

Given the following premises:

1. In the absence of ANY selection, fitness of individuals are at a theoretical maximum.
2. If a theoretical maximum fitness is achieved the effective population size is at a maximum.
3. If there is an allele that confers both increased fitness and decreased fitness (simultaneously) you have a genetic conflict (e.g. an allele that turns you green gives you good woodland camouflage but is un-attractive).
4. If a deleterious allele is linked to a beneficial allele this is similar to having a genetic conflict (mentioned in 3.).

The conclusion follows:

Strong linkage between beneficial alleles and deleterious alleles (Hill Robertson effects, i.e. 3. and 4.) prevents a theoretically maximum fitness to be attained (1.) and this a reduced effective population size (2.). But one which can be overcome by high levels of recombination and mating (since recombination can separate deleterious and beneficial alleles).

Hopefully this syllogism will answer your question.

Given the following premises:

1. In the absence of selection, fitness of individuals are at a theoretical maximum.
2. If a theoretical maximum fitness is achieved the effective population size is also at a maximum.
3. If there is an allele that confers both increased fitness and decreased fitness (simultaneously) you have a genetic conflict (e.g. an allele that turns you green gives you good woodland camouflage but is un-attractive).
4. If a deleterious allele is linked to a beneficial allele this is similar to having a genetic conflict (mentioned in 3.).

The conclusion follows:

Strong linkage between beneficial alleles and deleterious alleles (Hill Robertson effects, i.e. 3. and 4.) prevents a theoretically maximum fitness to be attained (1.) and thus a reduced effective population size (2.). Which can only be overcome by high levels of recombination and mating (i.e. by separating deleterious and beneficial alleles).